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Publications

Research Publications, Peer-Reviewed

  1. Mayoral R, Valverde AM, Llorente-Izquierdo C, González-Rodríguez A, Boscá L, Martín-Sanz P. Impairment of transforming growth factor beta signaling in caveolin-1- deficient hepatocytes: role in liver regeneration. J Biol Chem. 2010. Feb 5;285(6):3633-42. PMID: 19966340; PMCID: PMC2823504.
  2. Llorente-Izquierdo C, Mayoral R, Flores JM, García-Palencia P, Cucarella C, Boscá L, Casado M, Martín-Sanz P. Transgenic mice expressing cyclooxygenase-2 in hepatocytes reveal a minor contribution of this enzyme to chemical hepatocarcinogenesis. Am J Pathol. 2011 Mar;178(3):1361-73. PMID: 21356386; PMCID: PMC3069875. First-author.
  3. Agra Andrieu N, Motiño O, Mayoral R, Llorente-Izquierdo C, Fernández-Alvarez A, Boscá L, Casado M, Martín-Sanz P. Cyclooxygenase-2 is a target of microRNA-16 in human hepatoma cells: Role in proliferation and apoptosi PLoS One. 2012;7(11): e50935. PMID: 23226427; PMCID: PMC3511388.
  4. Fernández-Alvarez A#, Llorente-Izquierdo C#, Mayoral R, Agra N, Boscá L, Casado M, Martín-Sanz P. Evaluation of epigenetic modulation of Cyclooxygenase-2 as prognostic marker for hepatocellular carcinoma. Oncogenesis. 2012 Jul 9;1: e23. PMID: 23226427; PMCID: PMC3511388. #equal contribution. #First-author, equal contribution.
  5. Llorente-Izquierdo C, Mayoral R, Cucarella C, Grau C, Alvarez MS, Flores JM, García-Palencia P, Agra N, Castro-Sánchez L, Boscá L, Martín-Sanz P, Casado M. Progression of liver oncogenesis in the double transgenic mice c-myc/TGF-α is not enhanced by cyclooxygenase-2 expression. Prostaglandins Other Lipid Mediat. 2013 Oct; 106:106-15. PMID: 23579063. First-author.
  6. Castro-Sánchez L, Agra N, Llorente-Izquierdo C, Motiño O, Casado M, Boscá L, Martín-Sanz P. Regulation of 15-hydroxyprostaglandin dehydrogenase expression in hepatocellular carcinoma. Int J Biochem Cell Biol. 2013 Nov;45(11):2501-11. PMID: 23954207.
  7. Mayoral R, Osborn O, McNelis J, Johnson AM, Oh DY, Izquierdo CL, Chung H, Li P, Traves PG, Bandyopadhyay G, Pessentheiner AR, Ofrecio JM, Cook JR, Qiang L, Accili D, Olefsky JM. Adipocyte SIRT1 knockout promotes PPARγ activity, adipogenesis and insulin sensitivity in chronic-HFD and obesity. Mol Meta 2015 Mar 5;4(5):378-91. PMID: 25973386; PMCID: PMC4421024.
  8. Wang L, Fouts DE, Stärkel P, Hartmann P, Chen P, Llorente C, DePew J, Moncera K, Ho SB, Brenner DA, Hooper LV, Schnabl B. Intestinal REG3 Lectins Protect against Alcoholic Steatohepatitis by Reducing Mucosa-Associated Microbiota and Preventing Bacterial Translocation. Cell Host Microbe. 2016 Feb 10;19(2):227-39. PMID: 26867181; PMCID: PMC4786170.
  9. Hartmann P, Seebauer CT, Mazagova M, Horvath A, Wang L, Llorente C, Varki NM, Brandl K, Ho SB, Schnabl B. Deficiency of intestinal mucin-2 protects mice from diet-induced fatty liver disease and obesity. Am J Physiol Gastrointest Liver Physiol. 2016 Mar 1;310(5): G310-22. PMID: 26702135; PMCID: PMC4773827.
  10. Inamine T, Yang AM, Wang L, Lee KC, Llorente C, Schnabl B. Genetic Loss of Immunoglobulin A Does Not Influence Development of Alcoholic Steatohepatitis in Mice. Alcohol Clin Exp Res. 2016 Dec;40(12):2604-261 PMID: 27739086; PMCID: PMC5133156.
  11. Yang AM, Inamine T, Hochrath K, Chen P, Wang L, Llorente C, Bluemel S, Hartmann P, Xu J, Koyama Y, Kisseleva T, Torralba MG, Moncera K, Beeri K, Chen CS, Freese K, Hellerbrand C, Lee SM, Hoffman HM, Mehal WZ, Garcia-Tsao G, Mutlu EA, Keshavarzian A, Brown GD, Ho SB, Bataller R, Stärkel P, Fouts DE, Schnabl B. Intestinal Fungi Contribute to Development of Alcoholic Liver Disease. J Clin Invest. 2017 Jun 30;127(7):2829-2841. PMID: 28530644; PMCID: PMC5490775.
  12. Llorente C, Jepsen P, Inamine T, Wang L, Bluemel S, Wang HJ, Loomba R, Bajaj JS, Schubert ML, Sikaroodi M, Gillevet PM, Xu J, Kisseleva T, Ho SB, DePew J, Du X, Sørensen HT, Vilstrup H, Nelson KE, Brenner DA, Fouts DE, Schnabl B. Gastric Acid Suppression Promotes Alcoholic Liver Disease by Inducing Overgrowth of Intestinal Enterococcus. Nat Commun. 2017 Oct 16;8(1):83 PMID: 29038503; PMCID: PMC5643518. First-author.
  13. Khanova E, Wu R, Wang W, Yan R, Chen Y, French SW, Llorente C, Pan SQ, Yang Q, Li Y, Lazaro R, Ansong C, Smith RD, Bataller R, Morgan T, Schnabl B, Tsukamoto H. Pyroptosis by caspase11/4-gardening-D pathway in alcoholic hepatitis in mice and patient Hepatology. 2018 May;67(5):1737-1753. PMID: 29108122; PMCID: PMC5906140.
  14. Hartmann P, Hochrath K, Horvath A, Chen P, Seebauer CT, Llorente C, Wang L, Alnouti Y, Fouts DE, Stärkel P, Loomba R, Coulter S, Liddle C, Yu RT, Ling L, Rossi SJ, DePaoli AM, Downes M, Evans RM, Brenner DA, Schnabl B. Modulation of the intestinal bile acid-FXR-FGF15 axis improves alcoholic liver disease in mice.  2018 Jun;67(6):2150-2166. PMID: 29159825; PMCID: PMC5962369.
  15. Calvente CJ, Tameda M, Johnson CD, Del Pilar H, Lin YC, Adronikou N, De Mollerat Du Jeu X, Llorente C, Boyer J, Feldstein AE. Neutrophils contribute to spontaneous resolution of liver inflammation and fibrosis via microRNA-223. J Clin Invest. 2019 Jul 11; 130:4091-4109. PMID: 31295147; PMCID: PMC6763256.
  16. Duan Y#, Llorente C#, Lang S, Brandl K, Chu H, Jiang L, White RC, Clarke TH, Nguyen K, Torralba M, Shao Y, Liu J, Hernandez-Morales A, Lessor L, Rahman IR, Miyamoto Y, Ly M, Gao B, Sun W, Kiesel R, Hutmacher F, Lee S, Ventura-Cots M, Bosques-Padilla F, Verna EC, Abraldes JG, Brown RS Jr, Vargas V, Altamirano J, Caballería J, Shawcross DL, Ho SB, Louvet A, Lucey MR, Mathurin P, Garcia- Tsao G, Bataller R, Tu XM, Eckmann L, van der Donk WA, Young R, Lawley TD, Stärkel P, Pride D, Fouts DE, Schnabl B. Bacteriophage targeting of gut bacterium attenuates alcoholic liver disease. 2019 Nov;575(7783):505-511. PMID: 31723265; PMCID: PMC6872939. # First-author, equal contribution.
  17. Chu H, Duan Y, Lang S, Jiang L, Wang Y, Llorente C, Liu J, Mogavero S, Bosques-Padilla F, Abraldes JG, Vargas V, Tu XM, Yang L, Hou X, Hube B, Stärkel P, Schnabl B. The Candida albicans exotoxin Candidalysin promotes alcohol-associated liver disease. J Hepatol. 2020 Mar;72(3):391-400. PMID: 31606552; PMCID: PMC7031049.
  18. Zhou R, Llorente C, Cao J, Gao B, Duan Y, Jiang L, Wang Y, Kumar V, Stärkel P, Bode L, Fan X, Schnabl B. Deficiency of Intestinal α1-2-Fucosylation Exacerbates Ethanol-Induced Liver Disease in Mice. Alcohol Clin Exp Res. 2020 Sep;44(9):1842-1851. PMID: 32628772; PMCID: PMC7808344.
  19. Zhou R#, Llorente C#, Cao J, Zaramela LS, Zeng S, Gao B, Li SZ, Welch RD, Huang FQ, Qi LW, Pan C, Huang Y, Zhou P, Wang L, Liu EH, Yu RT, Downes M, Evans RM, Goglin K, Fouts DE, Brenner DA, Bode L, Fan X, Zengler K, Schnabl B. Intestinal a-1-2-fucosylation contributes to obesity and steatohepatitis in mice. Cell Mol Gastroenterol Hepat 2021 Feb 22;12(1):293-320. PMID: 33631374; PMCID: PMC8166943. # First-author, equal contribution.
  20. Cabré N, Duan Y, Llorente C, Conrad M, Stern P, Yamashita D, Schnabl B. Colesevelam Reduces Ethanol-Induced Liver Steatosis in Humanized Gnotobiotic Mice. Cells 2021; 10:1496. PMID: 34198609; PMCID: PMC8232222.
  21. Duan Y, Chu H, Brandl K, Jiang L, Zeng S, Meshgin N, Papachristoforou E, Argemi J, Mendes BG, Wang Y, Su H, Sun W, Llorente C, Hendrikx T, Liu X, Hosseini M, Kisseleva T, Brenner DA, Bataller R, Ramachandran P, Karin M, Fu W, Schnabl B. CRIg on liver macrophages clear pathobionts and protects against alcoholic liver disease. Nat Commun. 2021; 2(1):7172. PMID: 34887405; PMCID: PMC8660815.
  22. Murillo-Saich J, Coras R, Meyer R, Llorente C, Lane NE, Guma M. Synovial tissue metabolomic profiling reveals biomarkers of synovial inflammation in patients with osteoarthritis. Osteoarthr Cartil Open. 2022 Jul 9;4(3):100295. doi: 10.1016/j.ocarto.2022.100295. eCollection 2022 Sep. PMID: 36474936; PMCID: PMC9718344.
  23. Zeng S, Hartmann P, Park M, Duan Y, Lang S, Llorente C, Wang Y, Cabré N, Fouts DE, Bacher P, Jung WH, Stärkel P, Schnabl B. Malassezia restricta promotes alcohol-induced liver injury. Hepatol Commun. 2023 Jan 27;7(2):e0029. PMID: 36706195; PMCID: PMC9988279.
  24. Cabré N, Hartmann P, Llorente C, Kouno T, Wang Y, Zeng S, Kim HY, Zhang X, Kisseleva T, Iyer S, Kudumala S, Schnabl B. IgY antibodies against cytolysin reduce ethanol-induced liver disease in mice. 2023 Jul 1;78(1):295-306.  PMID: 36811393; PMCID: PMC10293100.
  25. Hsu CL, Wang Y, Duan Y, Chu H, Hartmann P, Llorente C, Zhou R, Schnabl B. Differences in Bacterial Translocation and Liver Injury in Ethanol Versus Diet-Induced Liver Disease. Dig Dis Sci. 2023 Jul;68(7):3059-3069. PMID: 36807831; PMCID: PMC10313731.
  26. Kouno T, Zeng S, Wang Y, Duan Y, Lang S, Gao B, Hartmann P, Cabré N, Llorente C, Galbert C, Emond P, Sokol H, James M, Chao CC, Gao JR, Perreault M, Hava DL, Schnabl B. Engineered bacteria producing aryl-hydrocarbon receptor agonists protect against ethanol-induced liver disease in mice. Alcohol Clin Exp Res. 2023 May;47(5):856-867. PMID: 36871955; PMCID: PMC10795770.
  27. Fuster D, Llorente C, Garcia-Calvo X, Bolao F, Zuluaga P, Hernández-Rubio A, Casado-Carbajo J, Leis A, Muga R. Fungal plasma biomarkers in patients admitted for inpatient treatment of alcohol use disorder. Clin Exp Res (Hoboken). 2023 Aug;47(8):1582-1589. PMID: 37364901; PMCID: PMC10984271.
  28. Hernandez J, Llorente C; Ma S, Saptarshi S, Ghosh P, Schnabl B, Huang W J.M. The arginine methyltransferase PRMT5 promotes mucosal defense in the intestine. Life Science Alliance. 2023 Sep 4;6(11):e202302026. PMID: 37666668; PMCID: PMC10477432.
  29. Llorente C. Isolation of Myenteric and Submucosal Plexus from Mouse Gastrointestinal Tract and Subsequent Co-Culture with Small Intestinal Organoids. Cells. 2024 May 10;13(10):815. PMID: 38786037; PMCID: PMC11120043. Corresponding author.
  30. Zhang P, Liu J, Lee A, Tsaur I, Ohira M, Duong V, Vo N, Watari K, Su H, Kim JY, Gu L, Zhu M, Shalapour S, Hosseini M, Bandyopadhyay G, Zeng S, Llorente C, Zhao HN, Lamichhane S, Mohan S, Dorrestein PC, Olefsky JM, Schnabl B, Soroosh P, Karin M. IL-22 resolves MASLD via enterocyte STAT3 restoration of diet-perturbed intestinal homeostasis. Cell Metab. 2024 Oct 1;36(10):2341-2354.e6. PMID: 39317186.
  31. Fujiki J, Nakamura T, Kreimeyer H, Llorente C, Fouts DE, Schnabl B. Insertion sequence-mediated phage resistance contributes to attenuated colonization of cytolytic Enterococcus faecalisvariants in the gut. Microbiol Spectr. 2025 Apr 15;13(5):e0330324. doi: 10.1128/spectrum.03303-24. Epub ahead of print. PMID: 40231830; PMCID: PMC12054073.
  32. Cabré N, Fondevila MF, Wei W, Yamazaki T, Tonetti FR, Eguileor A, Garcia-Carbonell R, Meijnikman AS, Miyamoto Y, Mayo S, Wang Y, Zhang X, Trimbuch T, Lehnardt S, Eckmann L, Fouts DE, Llorente C, Tsukamoto H, Stärkel P, Schnabl B. Activation of intestinal endogenous retroviruses by alcohol exacerbates liver disease. J Clin Invest. 2025 May 13;135(13):e188541. doi: 10.1172/JCI188541. PMID: 40359032; PMCID: PMC12208555.
  33. Llorente C*#, Raya Tonetti* F, Bruellman R, Brea R, Pell N, Hartmann P, Maccioni L, Han H, Cabré N, Liu J, Eguileor A, Fondevila MF, Meijnikman AS, Hsu CL, Alghafri A, Zhou R, Gao B, Duan Y, Zhang P, Febbraio MA, Taniguchi K, Newberry RD, Fouts DE, Brenner DA, Stärkel P, Karin M#, Schnabl B#. mAChR4 suppresses liver disease via GAP-induced antimicrobial immunity. 2025 Oct;646(8083):180-189. doi: 10.1038/s41586-025-09395-z. Epub 2025 Aug 20. PMID: 40836099. *First-author. #Corresponding author.
  34. Hsu CL, Shukla S, Freund L, Chou AC, Yang Y, Bruellman R, Raya Tonetti F, Cabré N, Mayo S, Lim HG, Magallan V, Cordell BJ, Lang S, Demir M, Stärkel P, Llorente C, Palsson BO, Mandyam C, Boland BS, Hohmann E, Schnabl B. Gut microbial ethanol metabolism contributes to auto-brewery syndrome in an observational cohort. Nat Microbiol. 2026 Feb;11(2):415-428. doi: 10.1038/s41564-025-02225-y. Epub 2026 Jan 8. PMID: 41507585.
  35. Fondevila MF, Kreimeyer H, Hsu CL, Tamargo-Azpilicueta J, Day LZ, Gritsenko M, Attah K, Cabré N, Harberts A, Tonetti FR, Yang Y, Yamazaki T, Schöler D, Eguileor A, Blasio C, Meijnikman AS, Zhang X, Garcia-Carbonell R, Hook V, Zhou E, Sun Z, Jacobs JM, Turner JR, Llorente C; AlcHepNet Investigators; Glass CK, Stärkel P, Reinheckel T, Diaz-Moreno I, Gonzalez DJ, Schnabl B. Macrophage-derived cathepsin B disrupts intestinal tight junctions through occludin degradation and promotes alcohol-associated liver disease. J Hepatol. 2026 Feb 5:S0168-8278(26)00026-7. doi: 10.1016/j.jhep.2026.01.013. Epub ahead of print. PMID: 41654223.
  36. Raya Tonetti F, Han H, Freund L, Cabre N, Fondevila MF, Eguileor A, Mayo S, Stärkel P, Hsu CL, Llorente C. Parallel regulation of goblet cell-associated antigen passages by reduced microbial sensing and mAChR4 signaling in Muc2 deficiency prevents ethanol-induced liver injury. Hepatology. 2026 Mar 27. doi: 10.1097/HEP.0000000000001744. Epub ahead of print. PMID: 41894257. Corresponding author.
  37. Kreimeyer H, Fondevila MF, Harberts A, Raya Tonetti F, Schöler D, Münte E, Hartmann P, Llorente C, Hube B, LeibundGut-Landmann S, Stärkel P; AlcHepNet Investigators; Schnabl B. Peptide YY reduces cytotoxicity of Candida albicans in alcohol-associated liver disease. JHEP Rep. 2026 May;8(5):101718. doi: 10.1016/j.jhepr.2025.101718. Epub 2025 Dec 23. PMID: 41996830; PMCID: PMC13101792.
  38. Davids M, Herrema H, K Groen A, Galenkamp H, Zwinderman A, Palmu J, Havulinna A, Niiranen T, Knight R, Acherman Y, Franken R, Verheij J, Dukas M; Dutch Liver Pathology Panel; Bajaj J, Llorente C, Schnabl B, Nieuwdorp M, Meijnikman A. Lactic acid bacteria and endogenous ethanol mediate proton pump inhibitor-associated MASLD: a multicohort cross-sectional mediation analysis. Gut Microbes. 2026 Dec 31;18(1):2664712. doi: 10.1080/19490976.2026.2664712. Epub 2026 May 3. PMID: 42070114; PMCID: PMC13154942.
  39. Raya Tonetti F*#, Han H, Bajaj JS, Schubert ML, Sikaroodi M, Gillevet PM, Lang S, Demir M, Rahman IR, van der Donk WA, InTeam Consortium, Hsu CL, Llorente C#*. Non-Absorbable Antibiotics Worsen Alcohol-Associated Liver Disease in Gastric Acid-Suppressed Mice. Gut microbes, Resubmitted, Accepted. *First-author. #Corresponding author.

Reviews, Editorials and Conference Summaries, peer-reviewed

  1. Llorente C, Schnabl B. The gut microbiota and liver disease. Cell Mol Gastroenterol 2015 May 1;1(3):275-284. PMID: 26090511; PMCID: PMC4467911. First-author.
  2. Wang L, Llorente C, Hartmann P, Yang AM, Chen P, Schnabl B. Methods to determine intestinal permeability and bacterial translocation during liver disease. J Immunol Methods. 2015 Jun; 421:44-53. PMID: 25595554; PMCID: PMC4451427.
  3. Llorente C, Schnabl B. Fast-Track Clearance of Bacteria from Liver. Cell Host Microbe. 2016 Jul 13;20(1):1- PMID: 27414492. First-author.
  4. Bruellman R, Llorente C. A Perspective of intestinal immune-microbiome interactions in alcohol-associated liver disease. Int J Biol Sci. 2021;17(1):307-327. PMID: 33390852; PMCID: PMC7757023. Corresponding author.
  5. Llorente C, Rungratanawanich, Liangpunsakul S. Intestinal dendritic cells, gatekeepers preventing ethanol-1 induced liver disease. 2023 Jan 3. PMID: 36930746. First-author.
  6. Raya Tonetti F, Eguileor A, Mrdjen M, Pathak V, Travers J, Nagy LE, Llorente C. Gut-liver axis: Recent concepts in pathophysiology in alcohol-associated liver disease. Hepatology. 2023 Sep 1;78(3):697-701. PMID: 36930746; PMCID: PMC10315410. Corresponding author.
  7. Llorente C. The Imperative for Innovative Enteric Nervous System-Intestinal Organoid Co-Culture Models: Transforming GI Disease Modeling and Treatment. Cells. 2024 May 10;13(10):820. PMID: 38786042; PMCID: PMC11119846. First-author. Corresponding author.
  8. Llorente C. Commentary on the gut microbiome in alcohol use disorder and alcohol-associated liver disease. Alcohol Clin Exp Res (Hoboken). 2024 Aug;48(8):1466-1468. doi: 10.1111/acer.15382. Epub 2024 Jun 2. PMID: 38825713; PMCID: PMC11305903. First-author. Corresponding author.
  9. Tonetti FR, Eguileor A, Llorente C. Goblet cells: guardians of gut immunity and their role in gastrointestinal diseases. eGastroenterology. 2024 Oct;2(3):e100098. doi: 10.1136/egastro-2024-100098. PMID: 39524932; PMCID: PMC11542612. Corresponding author.
  10. Kreimeyer H, Llorente C, Schnabl B. Influence of Alcohol on the Intestinal Immune System. Alcohol Res. 2025 Mar 14;45(1):03. doi: 10.35946/arcr.v45.1.03. PMID: 40151622; PMCID: PMC11913448.
  11. Tschann MM, Vachharajani V, Redmond EM, Hoisington A, Cohen SE, New-Aaron M, Llorente C, Paloczi J, Keating CR, Rungratanawanich W, Burnham EL, Callaci JJ, Raju P, Zhong W, Mandal A, Zimmerly JR, Nuncio ASP, Mandrekar P, McCullough RL, McMahan RH, Wyatt TA, Yeligar SM, Kovacs EJ, Choudhry MA. New Developments on the Effects of Alcohol Use on Immunity, Inflammation and Organ Function: A Summary of the 2024 Alcohol and Immunology Research Interest Group (AIRIG) Meeting. 2025 Apr 21:S0741-8329(25)00056-4. doi: 10.1016/j.alcohol.2025.04.002. Epub ahead of print. PMID: 40267994.
  12. Melamed E, Rungratanawanich W, Liangpunsakul S, Maki KA, McCullough RL#, Llorente C#. Alcohol, aging, and the gut microbiome: Intersections of immunity, barrier dysfunction, and disease. 2025 Nov;128:1-12. doi: 10.1016/j.alcohol.2025.07.001. Epub 2025 Jul 9. PMID: 40645301. #Corresponding author
  13. Jiang Y, Llorente C. YBX1/CD36 positive feedback loop: novel molecular target for MASLD treatment. Int J Biol Sci. 2025 Aug 11;21(12):5164-5166. doi: 10.7150/ijbs.121866. PMID: 40959287; PMCID: PMC12435216. Corresponding author.
  14. Gao B, Arab JP, Liangpunsakul S, Ding WX, Szabo G, Mehal W, Wang H, He Y, Stärkel P, Llorente C, Schnabl B, Hwang S, Vandermerwe S, Gao Y, Zheng MH, Kim W, George J, Bataller R, Loomba R, Leggio L, Tacke F. Metabolic dysfunction and alcohol-associated liver disease (MetALD). eGastroenterology. 2025 Dec 25;3(4):e100319. doi: 10.1136/egastro-2025-100319. PMID: 41459093; PMCID: PMC12742176.
  15. Jiang Y, Llorente C. Gut-Liver Axis Molecular Mechanisms in Alcohol-Associated Liver Disease. Alcohol. 2026 Apr 4:S0741-8329(26)00188-6. doi: 10.1016/j.alcohol.2026.04.001. Epub ahead of print. PMID: 41942011. Corresponding author.
  16. Blinchevsky N, Weed M, Liangpunsakul S, Llorente C, Melamed E. Vagus Nerve Stimulation as an Emerging Therapeutic Strategy for Alcohol Use Disorder. Alcohol. [Accepted].
  17. Llorente C. Intestinal neutral ceramidase, microbial metabolites and epithelial fucosylation in MASH. [Accepted]. Estimated submission date: June 2026. Alcohol. Invited editorial. Accepted First-author. Corresponding author.

Other

  1. Llorente C#, Karin M#. Alcohol Blocks Gut Immune Gatekeepers, Compromising Defense Against Bacteria in ALD. Behind the Paper. Nature. Corresponding author.

Complete List of Published Work in MyBibliography: 

National Library of Medicine - C. Llorente